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Awesome Pan-Cancer Phosphoproteomics

Pan-cancer phosphoproteomics can identify thousands of sites — but without knowing which kinase, which pathway, and which drug, a catalog is not a map.

Mass spectrometry can now quantify over 100,000 phosphorylation sites across tumor types. Yet fewer than 5% of these sites have a known upstream kinase, and fewer than 3% have established functional roles. The gap between measurement and understanding defines this field.

This site is the companion to the awesome list. While the list provides a curated index of tools, databases, and studies, this site goes deeper — with concept explainers, method comparisons, paper analyses, and synthesis essays that connect the pieces.

The Central Problem

What we can measure What we know
~240,000 human phosphosites cataloged <5% have a known upstream kinase
110,274 sites quantified across 10 CPTAC cancers <3% have established functional roles
518 kinases in the human kinome 130+ kinases have zero known substrates

The annotation bottleneck is not just a data problem — it shapes which biological questions can be asked and which therapeutic strategies can be pursued.

How This Site Is Organized

  • Concepts — Phosphorylation biology, the kinase-substrate inference problem, and how phosphoproteomics complements genomics.
  • Methods — Detailed guides for each tool category: identification, kinase inference, pathway reconstruction, functional scoring, pan-cancer integration, and therapeutic response.
  • Deep Reads — Honest analyses of landmark papers.
  • Benchmarks — The state of evaluation and what metrics actually matter.
  • Datasets — Key datasets organized by what analyses they support.
  • Timeline — Interactive literature timeline showing how the field evolved.
  • Explorer — Interactive maps: a method taxonomy and a kinase-substrate-pathway-drug biology network.
  • Synthesis — Cross-cutting essays on the annotation bottleneck and the path from phosphosites to signaling circuits.
  • Glossary — Key terms and definitions.

Key Numbers

Statistic Value Source
Human phosphosites cataloged ~240,000 PhosphoSitePlus
Sites with known upstream kinase <5% Community estimate
Sites with known function <3% Ochoa et al. 2020
Ser/Thr kinases with motif data 303 KinaseLibrary (2023)
CPTAC cancer types profiled 10+ Geffen et al. 2023
Kinase inhibitors FDA-approved 70+ As of 2024
Phosphosites per typical experiment 10,000–40,000 DIA-NN / MaxQuant